The Glasgow Warriors player has missed the last three weeks due to a shoulder injury but he has come through training unscathed and will feature at centre.
Captain Greig Laidlaw will partner stand-off Finn Russell for the clash at Murrayfield as Vern Cotter's men aim to build on their World Cup campaign.
John Barclay will make his first Six Nations appearance in four years at blindside with John Hardie and
David Denton while brothers Jonny and Richie Gray starting their tenth test match together.
Tommy Seymour, Sean Maitland and Stuart Hogg all start against Eddie Jones' men in Edinburgh with Cotter calling on his team to believe they can win the championship.
Scottish Rugby
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@Scotlandteam
YOUR @Scotlandteam to face England in Saturday's @rbs_6_nations clash at BT Murrayfield (kick-off 4.50pm)
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